In evolutionary biology, a spandrel is a phenotypic trait that is a byproduct of the evolution of some other characteristic, rather than a direct product of adaptive selection. Stephen Jay Gould and Richard Lewontin brought the term into biology in their 1979 paper "The Spandrels of San Marco and the Panglossian Paradigm: A Critique of the Adaptationist Programme".
In evolutionary biology, a spandrel is a phenotypic trait that is a byproduct of the evolution of some other characteristic, rather than a direct product of adaptive selection. Stephen Jay Gould and Richard Lewontin brought the term into biology in their 1979 paper "The Spandrels of San Marco and the Panglossian Paradigm: A Critique of the Adaptationist Programme".
Source: Wikipedia "Spandrel (biology)" · CC BY-SA 4.0
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